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Susan Cooper (Melissa McCarthy) is an unassuming, desk-bound CIA analyst guiding her partner Agent Bradley Fine (Jude Law) on a mission to Bulgaria from the CIA office in the US, using a radio connected to his earpiece. After witnessing Fine accidentally killing his target without finding the suitcase nuke bomb they are after, Susan guides him on his escape back to the United States. Meanwhile, the agency learns that the target's daughter Rayna Boyanov (Rose Byrne) might know the location of her father's device, so they send Fine to infiltrate her home. However, Fine is apparently shot dead by Rayna, which enrages Susan, who also discovers that Rayna had known the identities of all the agency's top agents, including Fine and Rick Ford (Jason Statham). After Fine's funeral, Susan volunteers to become a field agent. Since she never was one before, her boss Elaine Crocker (Allison Janney) agrees as long as she does not get close to either Rayna or her contact, Italian playboy Sergio De Luca (Bobby Cannavale), as she could break her cover. Ford quits, disgusted with Susan being chosen over him for the assignment.
Susan is sent to Paris, where the agency places her to spy on De Luca. After unexpectedly finding Ford there also, she sees an assassin working for De Luca (Nargis Fakhri) exchanging his bag with a bomb; she is successful in warning him which leads to Ford throwing the bomb into the river where it explodes. Susan then follows the assassin, but he is killed during a fight with Susan when he falls off of a high rise building. Susan follows De Luca to Rome, where she meets her contact Aldo (Peter Serafinowicz), a clumsy informant who makes inappropriate advances towards Susan. She soon decides to break her cover by going inside a casino just to retrieve more intelligence on De Luca. She instead meets Rayna, and quickly becomes part of Rayna's inner circle after successfully preventing Rayna's assassination. Gaining Rayna's trust, the two use her private plane to go to Budapest. Along the way, the pilot tries to kill Rayna, but Susan is able to subdue him and take control of the plane in time before it crashes onto the Alps. When Rayna believes Susan to be an undercover spy, Susan convinces her that her father secretly hired Susan to be Rayna's bodyguard. In Budapest, the two encounter Susan's best friend Nancy (Miranda Hart), who is sent by Crocker to make sure of Susan's condition, and Aldo. When a vehicle fires at them, Nancy tries to get Rayna to safety while Susan pursues the vehicle on a Vespa. Susan finds out that the shooter is CIA double agent Karen Walker (Morena Baccarin), who sold Rayna the names of the agents. As Walker is about to kill Susan, she is suddenly shot and killed by an unknown sniper.
That night, Susan and Nancy are set to witness Rayna meeting a contact who intends to purchase the nuclear device at a party. The contact turns out to be De Luca's associate, who Susan pursues on foot while Nancy gets arrested for assaulting 50 Cent on stage to distract Rayna from seeing the associate. Susan fights the assassin in a kitchen, exposing her true identity as a spy just before Fine, revealed to be still alive, throws a knife at the assassin's chest. He states he is working with Rayna and has become her lover. When Rayna has Susan and Aldo locked in a warehouse, Fine goes to Susan and reveals he shot Karen Walker and that he faked his death in order to gain intelligence on the device's location by posing as Rayna's lover. Eventually, Aldo and Susan cooperate to escape, with Susan joining Rayna and Fine at De Luca's mansion by posing as a double agent who's betraying her agency. A third party, led by Russian gangster Dudaev, arrives only to be shot dead by De Luca, who reveals his true motives of wanting to an even higher bidder. As De Luca tries to kill Rayna next, Ford spectacularly fails his rescue entrance, adistraction that allows Susan to save Rayna's life yet again. De Luca escapes with the device on his helicopter, wounding Fine in the process. S usan and Ford jump onto De Luca's helicopter, but Ford soon loses balance and falls down into the lake, leaving Susan to confront De Luca over the device. With the help of Nancy, Aldo, and 50 Cent (who appears to have a mutual sexual interest in Nancy) in another helicopter, Susan is able to kill De Luca as both he and the device fall down to the lake with less success than Ford.
In the aftermath of her successful mission, Crocker arrives to recover the device, as well as promoting Susan to field agent, which she happily accepts. 50 Cent gloats over his involvement in taking down the terrorists, going as far as mocking fellow rapper/rival Kanye West for never accomplishing such a thing. Fine is treated for his wounds, and Ford takes off on a speed boat to seaside destinations, unaware he's on a lake. Rayna is arrested, but not before she and Susan jokingly exchange profane insults subtly implying that Rayna has grown to like Susan. Aldo re-introduces himself as Albert, an undercover British MI6 agent who invites Susan to dinner should she ever find herself in London, which Susan accepts. Fine then asks Susan to dinner but she declines, deciding to have a "girl's night" with Nancy.
Before the closing credits roll, Susan is shown screaming after waking up and realizing that she has spent the night with Ford.
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